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324       The Rise of Rudists; Middle Cretaceous Facies in  Mexico and the Middle East


                     Ool;t. ~ (ool.d Dra in
                     land  bar
                     .",dil'  debris ~=~:;:g~~7~~~~~~~~~:;;
                     bioclastic


                          TOP  Of  T ... NINUL  au ..... y.  CLOSE  TO  E"'ST  F.ONT  Of  V ... LLES  P~TfORM
                          Mod.,ot.l,  kigh  ••• ,gy    HE ... R V"'LLES,  SLP,  MEX.








                     10UCO'io.hn.It,;:,0~:~~§~~~~~~~~~~~~§~~~=
                               1  ~
                     lobric;
                              __
                          RUD IST  BUILDUPS  Of  ... CTOP ... N  .... NK  WITH IN  EL  "'U'"  SHELF,  CEN.  MEXICO
                          ... tlle,.d  ci rculation  ond  en.rGY  Itrollll  a. Cono.(.o  IMP,  Me",)
              Open  lea  . id.                    IOkm
                                                        -----
                                                         Coprinid.  -----
             8·IOm

                                _--     &'I   ../'  t1 ......,.",.   (l\
                        --,.--~     ./"   rr  Touc:cu lo  and   .....".--!M:::!o~no=I."".,"'"o ____   _
                                   v   .,   \1  Clodophyllio  V   'to   V
                        IDE ... LlZED  IUILDUP  (T .... UlAR  REEf)  SHOWING  I I0f ... CIES  PROGUSSION  TO  SHO ... LlNG  W"'TER-
                        INNER  SHELf  OF  TEX ... S  CIl ... 10N   F,om  K.  To •• g ,  1959  modili.d  by  ....  Coogon
               Fig.XI-4. Facies in three rudist mounds on shelves or along gently dipping shelf margins in
               Texas and Mexico


               ids together in extreme backreef positions, e.g., in the EI  Abra of Mexico. Figure
               XI-3  shows  a  facies  profile  with  the  ecological  zonation  of  the  major  rudist
               groups and associated biota. Figure XI-4  is  an illustration of the composition of
               typical rudist mounds,
                  Rudists  differ  somewhat from  other common reefoid  organisms  in  the  geo-
               logic record. This is  true both in depositional environment and in habit of con-
               struction.
                  1.  Generally in the Middle Cretaceous (except for monopleurids) they inhab-
               ited clear-water, pure carbonate environments without clay contaminants. In the
               Late Cretaceous some of the radiolitids and hippuritids were also tolerant of the
               clayey bottom.
                  2.  They were somewhat tolerant of variable salinity and restricted circulation.
               The absence of brachiopods, ammonites, and echinoids from typical rudist facies
               indicates that the organism could withstand fluctuations in temperature, salinity,
               and oxygen beyond the limits  of normal  marine water.  Discovery of numerous
               diastems and hard grounds within caprinid mounds in Texas, and the  presence  of
               coal  directly  above  caprinid  (?)  biostromes  in  the  Middle  East, indicates  that
               some rudists could survive intermittent exposure.
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