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Physical, chemical, and tensile properties of cashmere, mohair, alpaca  117

           found increasing proportions of orthocortical cells with increasing fiber and cortical
           diameter. The reversion to mainly orthocortial cells under nutritional stress indicated
           that this was the basic cortical cell in wool. Single follicles produced varying propor-
           tions of all cortical cell types and organized them in a variety of ways within the fiber.
              Only one study investigating of the size of cortical cells in a range of rare animal
           fiber is known (McGregor and Quispe, 2017). There was a large relative range in
           the size of cortical cell length, diameter, and volume. Cortical cell diameter showed
           less variation (mean diameters 4.5e4.9 mm) than cell length (mean lengths
           42.5e57.3 mm). The relative shape of cortical cells was predictable as cortical cell
           length was strongly related to cortical cell diameter. The ratio length:diameter
           increased by 0.19/1 mm increase in cell length and declined by 0.03/1 mm increase
           in MFD. Generally the cortical dimensions of these animal fibers were similar with
           the exception that qiviut had longer cortical cells with a greater volume compared
           with cashmere, mohair, bison, and vicu~ na.
              Nutritional manipulation resulted in cashmere grown by goats with higher levels of
           nutrition having longer cortical cells compared with cashmere grown by goats in
           restricted nutrition treatments (50.8 vs. 34.5 mm) with greater diameter (4.57 vs.
                                        3
           4.14 mm), volume (274 vs. 170 mm ), and a higher length:diameter ratio (11.1 vs.
           8.4). These results indicate that the internal physical structure of cashmere fibers are
           not fixed and are influenced by nutritional status (McGregor and Liu, 2017).

           4.2.2.3  Medullation

           Fibers that have a hollow or a partially filled central canal running either as a contin-
           uous or in a fragmented form along their length are known as medullated fibers, and
           are present to some extent in the fleece of all animals. Some of these fibers have a
           chalky white appearance and are often referred to as “kemp.” Kemp fibers show other
           characteristics including being relatively shorter, coarser, more brittle and pigmented,
           and have flattened portions and sharp bends compared with normal fibers (Frazer
           Roberts, 1926). The medullated fibers are those in which the diameter of the medulla
           is less than 60% of the diameter of the fiber. Kemp fibers are medullated fibers in
           which the diameter of the medulla is 60% or more of the diameter of the fiber when
           viewed in longitudinal section. Kemp fibers have a different cross-section, which is
           oval or kidney shaped, compared with wool that approximates a circular cross-
           section. In mohair, Lupton et al. (1991) showed that a limiting thickness of about
           5 mm of solid keratin surrounds the medulla, that the limiting minimum medulla thick-
           ness was 5 mm, and generally the diameter of the medulla increases with increasing
           fiber diameter.
              All medullated fibers are contaminants in mohair, cashmere, alpaca, llama, vicu~ na,
           and yak fibers. The medullated fibers present as guard hairs and kemp can be removed
           by commercial dehairing. The content of medullated fibers in raw fibers is substantial:
           for combed Chinese cashmere 8%e15%; shorn cashmere fleeces 50%e75%; llama
           30%e70%; alpaca 5%e90%; Dromedry camel 40% (Frank et al., 2011; McGregor,
           2006a; Msahli et al., 2008). Good-quality greasy mohair has <1% medullated fiber
           and after processing the level is generally <0.3% (Hunter, 1993). For high-quality
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