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MASS EXTINCTIONS AND BIODIVERSITY LOSS  181


             single islands (e.g. the dodo) or in extreme    3  Investigate one of the “other” mass extinc-
             conditions (e.g. the Great auk). Perhaps more      tions not covered in detail here: end-Ordo-
             widespread species such as pigeons, sparrows       vician, Late Devonian and end-Triassic.
             and chickens will survive such depredations?    4  Calculate the relative magnitudes of the

             But recall the Passenger pigeon – it should        big five events from Jack Sepkoski’s data-
             have been immune to extinction. The other          base of fossil genera, either through http://
             point is to query whether it is right to extrap-   strata.ummp.lsa.umich.edu/jack/ or http://

             olate the figures from bird and mammal              geology.isu.edu/FossilPlot/.
             extinctions to the rest of life. Species of birds   5  Why is the current loss of species on
             and mammal are short-lived (i.e. they evolve       Earth sometimes termed the “sixth
             fast), and perhaps their extinction rates are      extinction”?
             not appropriate for insects and plants, for
             example.
               The jury is still out on modern extinction.    Further reading
             It is clear that surging human population and
             increasing tension between development and      Benton, M.J. 2003.  When Life Nearly Died. W.W.
                                                               Norton, New York.
             ecology put pressure on natural habitats and    Benton, M.J. &  Twitchett, R.J. 2003. How to kill
             on species. Plants and animals are dying out      (almost) all life: the end-Permian extinction event.
             faster now than at times in the past when the     Trends in Ecology and Evolution 18, 358–65.
             global human population was smaller. Pale-      Briggs, D.E.G. & Crowther, P.R. 2001. Palaeobiology,
             ontologists and ecologists have an important      A Synthesis, 2nd edn. Blackwell, Oxford, UK.
             job to do in seeking to understand just what    Erwin, D.H. 2006.  Extinction: How Life on Earth
             the threats are and how fast the modern           Nearly Ended 250 Million Years Ago. Princeton Uni-
             extinction is proceeding.                         versity Press, Princeton, NJ.
                                                             Gotelli, N.J. & Colwell, R.K. 2001. Quantifying biodi-
                                                               versity: procedures and pitfalls in the measurement
                                                               and comparison of species richness. Ecology Letters
              Review questions                                 4, 379–91.
                                                             Hallam, A. & Wignall, P.B. 1997. Mass Extinctions and
             1  How do paleontologists and other earth         their Aftermath. Oxford University Press, Oxford,
                scientists study mass extinctions? Carry       UK.
                out a census of papers about the Permo-      Hammer, Ø. & Harper, D.A.T. 2005.  Paleontological
                Triassic event published in the last year.     Data Analysis. Blackwell Publishing, Oxford, UK.

                Find the first 50 papers using any biblio-    Jablonski, D. 2005. Mass extinctions and macroevolu-
                graphic search tool, and classify them by      tion. Paleobiology 31, 192–210.
                broad theme (paleontology, stratigraphy,     Taylor, P. 2004. Extinctions in the History of Life. Cam-
                geochemistry, atmospheric modeling, vol-       bridge University Press, Cambridge, UK, 204 pp.
                canology), geographic region (perhaps by
                continents), sedimentary regime (marine,
                terrestrial) and key conclusion about the     References
                extinction model (eruption of Siberian       Alvarez, L.W., Alvarez, W., Asaro, F. & Michel, H.V.
                Traps, gas hydrate release, acid rain,         1980 Extraterrestrial cause for the Cretaceous-
                anoxia, meteorite impact). How are our         Tertiary extinction. Science 208, 1095–108.
                views perhaps biased by limited geo-         Bambach, R.K. 2006. Phanerozoic biodiversity mass
                graphic coverage, a major focus on marine      extinctions. Annual Review of Earth and Planetary
                rocks and dominant academic discipline?        Sciences 34, 127–55.

                Are these biases to be expected, and         Benton, M.J. 1995. Diversification and extinction in the
                why?                                           history of life. Science 268, 52–8.
             2  Is there any evidence that the media dis-    Hammer, Ø. & Harper, D.A.T. 2005.  Paleontological
                torts research agendas? Look at news           Data Analysis. Blackwell Publishing, Oxford, UK.
                stories about the KT event, and consider     Jablonski, D. 2005. Mass extinctions and macroevolu-
                                                               tion. Paleobiology 31, 192–210.
                the balance of reporting of different        Jin, Y.G., Wang, Y., Wang, W., Shang, Q.H., Cao, C.Q.
                aspects: do a census of the animal and         & Erwin D.H. 2000. Pattern of marine mass extinc-
                plant groups mentioned in the fi rst  50        tion near the Permian-Triassic boundary in South
                news reports you encounter.                    China. Science 289, 432–6.
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