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316  INTRODUCTION TO PALEOBIOLOGY AND THE FOSSIL RECORD





                                                          lophophore






                                                                collar


                                          muscular
                                          sphincter
                                                                                     parietal
                                                                                     muscles
                                                                                      gut
                                                                                      transverse
                                                                                      parietal
                                                                                      muscles
                                            metacel
                                           retractor
                                           muscle
                                                                                     tissue
                                                                                     cords



                                                                stolon
                                           (a)




                                                                                      lophophore

                                                frontal   retractor  operculum
                                                membrane  muscles                     anus
                                                                   parietal
                                                                   muscles
                                                                                      operculum
                                                                                      closing
                                                                                      muscle
                                                                                      metacel

                                                  ovary tissue  testis  gut  lateral
                                                       cord               pores
                                          (b)
                      Figure 12.15  Morphology of two living bryozoans: (a) a stenolaemate and (b) a gymnolaemate. (Based
                      on various sources.)



                      have modeled the feeding strategies of these      In general both the cryptostomes and fenes-
                      screw-shaped colonies and other fenestrates.    trates outstripped the trepostomes during the
                      Carboniferous fenestrate colonies usually had   Late Paleozoic, many of the fenestrates popu-
                      inward-facing zooids and probably drew          lating reef environments. Although both
                      water in through the top of the colony and      groups disappeared at the end of the Permian

                      flushed it out through the fenestrules at the    or soon after, they were still conspicuous
                      sides. On the other hand, Silurian colonies     members of the Late Permian benthos; both
                      had outward-facing zooids and sucked in         Fenestella and  Synocladia form large, vase-
                      water through the fenestrules, expelling it out   shaped colonies in the communities of the
                      of the open top of the colony.                  Zechstein reef complex in the north of England
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