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352  INTRODUCTION TO PALEOBIOLOGY AND THE FOSSIL RECORD










                                                                   Crioceras sp. 1  Crioceras sp. 2
                                   Dactylioceras  Normannites
                                                                                                       ?
                                                                                                     Turrilites

                                             Promicroceras
                                                                               Macroscaphites
                               Caloceras
                                                                   Crioceras sp. 3




                                                                   Scaphites
                               Ludwigia       Sigaloceras                     Oecoptychius      Lytocrioceras
                          (a)









                                   100 m                  100 m                 100 m









                                    oxygenated




                                                            anoxic               anoxic



                                  (b)
                      Figure 13.21  Life attitudes and buoyancy of the ammonites. (a) Supposed life orientations of a
                      selection of ammonite genera, with the center of gravity marked ×; the center of buoyancy is marked
                      with a dot and the extent of the body chamber is indicated with subparallel lines. (b) Relationship of
                      some ammonite morphotypes to water depth and the development of anoxia. (a, from Trueman, A.E.
                      1940. Q. J. Geol. Soc. Lond. 96; b, from Batt 1993.)


                      sacs and body outline is well known from        ing with the exoskeletons of the shelled ceph-
                      several localities in the Jurassic. In contrast,   alopods such as the nautiloids and ammonoids.
                      the skeletons of extinct belemnites are locally   The belemnite skeleton is relatively simple,
                      abundant in Jurassic and Cretaceous rocks.      consisting of three main parts: the bullet-
                      Belemnites had an internal skeleton, contrast-  shaped guard is solid and composed of radi-
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