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360  INTRODUCTION TO PALEOBIOLOGY AND THE FOSSIL RECORD


                                                                        Middle Cambrian Burgess Shale.  Nature  442,
                       Review questions                                 159–163.
                                                                      Christensen, W.K. 2000. Gradualistic evolution in
                      1  There has been some diffi culty identifying
                         the first mollusk. What are the key fea-        Belemnitella from the middle Campanian of Lower

                                                                        Saxony, NW Germany.  Bulletin of the Geological
                         tures of the phylum and how would they         Society of Denmark 47, 135–63.
                         be recognized in the fi rst mollusk?          Doyle, P. & MacDonald, D.I.M. 1993. Belemnite battle-
                      2  Many taxa that form part of the Early          fi elds. Lethaia 26, 65–80.
                         Cambrian biota are undoubtedly mol-          Erwin, D.H. 2007. Disparity: morphological pattern
                         lusks. Which mollusk groups are already        and developmental context.  Palaeontology  50,
                         present in the small shelly fauna?             57–73.
                      3  Theoretical morphospace is a useful tool     Fedonkin, M. & Waggoner, B.M. 1997. The Late Pre-
                         to investigate shell morphology. Some          cambrian fossil Kimberella is a mollusk-like bilate-
                         groups are more constrained in their           rian organism. Nature 388, 868–71.
                         developmental opportunities than others.     Harper, E.M. 2006. Disecting post-Palaeozoic arms
                         What advantages should univalved mol-          races.  Palaeogeography, Palaeoclimatology, Palaeo-
                                                                        ecology 232, 322–43.
                         lusks have over bivalved mollusks in a       Jacobs, D.K. & Landman, N.H. 1993.  Nautilus – a
                         quest to generate extreme morphotypes?         poor model for the function and behavior of ammo-
                      4  Belemnites seem an unlikely group to test      noids. Lethaia 26, 101–11.
                         models for microevolution. What condi-       Milsom, C. & Rigby, S. 2004.  Fossils at a Glance.
                         tions should be met in such tests of micro-    Blackwell Publishing, Oxford.
                         evolutionary hypotheses?                     Peel, J.S. 1991. Functional morphology, evolution and
                      5  The Mesozoic marine revolution (or arms        systematics of early Palaeozoic univalved molluscs.
                         race) was a complex ecological event that      Grønlands Geologiske Undersøgelse 161, 116 pp.
                         set the agenda for marine life in the        Raup, D.M. 1966. Geometric analysis of shell coiling:
                         Modern evolutionary fauna. How did             general problems.  Journal of Paleontology  40,
                         mollusks react to predation pressures?         1178–90.
                                                                      Reboulet, S., Giraud, F. & Proux, O. 2005. Ammonoid
                                                                        abundance variations related to changes in trophic
                       Further reading                                  conditions across the Oceanic Anoxic Event 1d
                                                                        (Latest Albian, SE France). Palaios 20, 121–41.
                      Clarkson, E.N.K. 1998. Invertebrate Palaeontology and
                                                                      Runnegar, B. & Pojeta, J. 1974. Molluscan phylogeny:
                        Evolution, 4th edn. Chapman and Hall, London.
                                                                        the palaeontological viewpoint. Science 186, 311–17.
                        (An excellent, more advanced text; clearly written
                                                                      Sigwart, J.W. & Sutton, M.D. 2007. Deep molluscan
                        and well illustrated.)
                                                                        phylogeny: synthesis of palaeontological and neon-
                      Lehmann, U. 1981. The Ammonites – their Life and their
                                                                        tological data.  Proceedings of the Royal Society  B
                        World. Cambridge University Press, Cambridge, UK.
                                                                        274, 2413–19.
                      Morton, J.E. 1967. Molluscs. Hutchinson, London.
                                                                      Stanley, S.M. 1970. Relation of shell form to life habits
                      Peel, J.S., Skelton, P.W. & House, M.R. 1985. Mollusca.
                                                                        of Bivalvia. Geological Society of America Memoir
                        In Murray, J.W. (ed.) Atlas of Invertebrate Macrofos-
                                                                        125, 296 pp.
                        sils. Longman, London. (A useful, mainly photo-
                                                                      Swan, A.R.H. 1990. Computer simulations of inverte-
                        graphic review of the group.)
                                                                        brate morphology. In Bruton, D.L. & Harper, D.A.T.
                      Pojeta, J. Jr., Runnegar, B., Peel, J.S. & Gordon, M. Jr.
                                                                        (eds)  Microcomputers in Palaeontology. Contribu-
                        1987. Phylum Mollusca.  In Boardman, R.S.,
                                                                        tions from the Palaeontological Museum, University
                        Cheetham, A.H. & Rowell, A.J. (eds) Fossil Inverte-
                                                                        of Oslo, Vol. 370. pp. 32–45. University of Oslo,
                        brates. Blackwell Scientific Publications, Oxford,

                                                                        Oslo.
                        UK, pp. 270–435. (A comprehensive, more advanced
                                                                      Vinther, J. & Nielsen, C. 2004. The Early Cambrian
                        text with emphasis on taxonomy; extravagantly
                                                                        Halkieria is a mollusc. Zoologica Scripta 34, 81–9.
                        illustrated.)
                                                                      Wagner, P.J. 1995. Diversity patterns among early gas-
                      Vermeij, G.J. 1987. Evolution and Escalation. An Eco-
                                                                        tropods: contrasting taxonomic and phylogenetic
                        logical History of Life. Princeton University Press,
                                                                        descriptions. Paleobiology 21, 410–39.
                        Princeton, NJ. (Visionary text.)
                                                                      Wani, R., Kase, T., Shigeta, Y. & De Ocampo, R. 2005.
                                                                        New look at ammonoid taphonomy, based on fi eld
                       References
                                                                        experiments with modern chambered nautilus.
                      Batt, R. 1993. Ammonite morphotypes as indicators of   Geology 33, 849–52.
                        oxygenation in a Cretaceous epicontinental sea.   Williamson, P.G. 1981. Palaeontological documentation
                        Lethaia 26, 49–63.                              of speciation in Cenozoic molluscs from Turkana
                      Caron, J.-B., Acheltema, A., Schander, A. & Rudkin, D.   basin. Nature 293, 140–2.
                        2006. A soft-bodied mollusc with radula from the
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