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DEUTEROSTOMES: ECHINODERMS AND HEMICHORDATES  423



                          Systems  series  stages  Graptolite  Formations  Lithology  fall            rise  Sequence  Environmental  features
                                                                   Relative
                                                                           -stratigraphic
                                                        and gaps in
                                                                   sea level
                                                                           units and
                                   biozones and subzones
                                                        sedimen-
                                                                           system tracts
                                                        tation
                             W.  Centrifugus                                          Upwelling
                                                                                       possibly
                                 Insectus                                             weakened
                                 Lapworthi  grandis   MOTOL                      HST
                                 Spiralis
                               Telychian  Tullbergi                          UNIT         4
                                 Griestoniensis
                                 Crispus              LITOHLAVY                  TST
                                 Turriculatus  hispanicus                            perturbations
                                                                                       Oceanic
                          Silurian  Llandovery  Linnaei  palmeus            sb  (LST)
                                 Sedgwickii
                                            tenuis
                               Aeronian  Convolutus  richteri                   HST
                                 Leptotheca
                                            paradenticul.
                                 Simulans
                                 Triangulatus-Pectinatus  ŽELKOVICE          UNIT         3  TST  Maximum
                                 Cyphus
                                                                                      upwelling
                               Rhuddanian  Vesiculosus  atavus              sb   LST
                                 Acuminatus
                                                                                       Anoxic
                                 Ascensus                                       HST    turnover
                                             shelly                                   deglaciation
                                 Persculptus                                 UNIT       2  TST
                                            Hirnantia
                                             fauna
                                                                                 LST  2nd glacial
                                                                            sb
                                          ?                                           maximum
                                                                                HST
                                                                                      interglacial
                                                                                 TST
                               Hirnantian             KOSOV                  UNIT         1  1st glacial



                          Ordovician  Ojsuensis                                 (LST)  maximum
                                                                                      Glaciation

                                   pre-Hirnantian shelly                   ? sb  HST  Climatically
                                                                                        warm
                                   fauna              KR. DVUR                         period
                                 Laticeps


                                                     storm sandstones,
                              shale, mudstone        siltstones and shale   limestone nodules
                                                                            and lenticular beds
                                                     interbeds
                                                     graded sandstone       alternating pale mudstone
                              black shale
                                                     and conglomerate       and black shale
                              sandy/ silty-micaceous  glaciomarine
                              laminite               diamictite
             Figure 15.30  Graptolite biostratigraphy of the Upper Ordovician–Lower Silurian strata of the
             Barrandian basin. HST, highstand systems tract; TST, transgressive systems tract; LST, lowstand systems
                            ˇ
             tract. (Based on Storch 2006.)
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