Page 445 - Introduction to Paleobiology and The Fossil Record
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432  INTRODUCTION TO PALEOBIOLOGY AND THE FOSSIL RECORD




                                           posterior  anterior             anterior
                                                                                    posterior




                                                basal cavity
                                                                                basal cavity
                                             Hertzina                          Furnishina
                                          (a)                           (b)
                                        anterior  posterior                   posterior  main cusp
                                                 main cusp                                anterior
                                     anterior
                                     blade              denticles                         denticles
                                                                                  basal
                                                              posterior           cavity
                                                              blade
                                             basal cavity                          Prioniodina
                                             Ozarkodina


                                          posterior
                                                                                        posterior
                                                     platform                            platform
                                         carina

                                                    basal cavity
                                                                                              secondary
                                                                                              platforms
                                                                           lateral
                                                                           process
                                            blade   inner side  outer side
                                                                                        blade
                                           anterior                                     anterior
                                             Polygnathus                           Amorphognathus
                                   (c)
                      Figure 16.3  Descriptive morphology of the main types of conodont elements: (a) protoconodont
                      Herzina (×40); (b) paraconodont Furnishina (×40); and (c) euconodonts Ozarkodina (×40), Prionodina
                      (×20), Polygnathus (×40) and Amorphognathus (×40). (Based on Armstrong & Brasier 2004.)



                      Paraconodonts are reported from the Mid         wide variety of blades and platforms in the
                      Cambrian; older records are doubtful. During    Mid to Late Ordovician. This great diversity
                      the Late Cambrian, simple conical eucon-        of forms was wiped out by the Late Ordovi-
                      odonts appeared. In the Early Ordovician,       cian mass extinction (see p. 169). Silurian
                      apparatuses with coniforms, and some with       faunas are less variable, mainly apparatuses
                      coniform and ramiform element types,            with ramiform and pectiniform elements. The
                      appeared. Conodont diversity peaked during      conodonts again radiated during the Late
                      the Mid Ordovician, with a global maximum       Devonian, with specialized ramiform and pec-
                      of over 60 genera. During this interval of      tiniform elements; over 1000 conodont taxa
                      experimentation, there was a huge diversity     have been named from the Upper Devonian.
                      of apparatus patterns never again matched;      Carboniferous conodonts (Fig. 16.5a) were
                      later apparatuses are relatively uniform,       characterized by a lack of coniform elements,
                      perhaps indicating stabilization of feeding     together with pectiniform elements in the P
                      modes. Pectiniform elements were common         apparatus position, whereas ramiform ele-
                      from the Early Ordovician, together with a      ments occupied the M and S positions (see
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