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200                                   Algae: Anatomy, Biochemistry, and Biotechnology

                  Irradiance (Flux Density)
                                                    2
                    1Wm   22  ¼ 10 24  Wcm 22  ¼ 6.83   10 lux (lx) at 555 nm
                    1lmm  22  ¼ 1lx ¼ 10 24  lm cm 22
                    1 photon at 400 nm ¼ 4.96   10 219  J
                    1Wm   22  at 400 nm ¼ 1Jm 22  sec 21  ¼ 3.3 mmol m 22 21  ¼ 3.3 mEinsten m 22  sec 21
                                                                 s
                    1 photon at 500 nm ¼ 3.97   10 219  J
                    1Wm   22  at 500 nm ¼ 1Jm 22  sec 21  ¼ 4.2 mmol m 22  sec 21  ¼ 4.2 mEinsten m 22  sec 21
                    1 photon at 600 nm ¼ 3.31   10 219  J
                    1Wm   22  at 600 nm ¼ 1Jm 22  sec 21  ¼ 5.0 mmol m 22  sec 21  ¼ 5.0 mEinsten m 22  sec 21
                    1 photon at 700 nm ¼ 2.83   10 219  J
                    1Wm   22  at 700 nm ¼ 1Jm 22  sec 21  ¼ 5.9 mmol m 22  sec 21  ¼ 5.9 mEinsten m 22  sec 21


                  Radiance
                       22  21           2    22  21                    22
                  1Wm     sr  ¼ 6.83   10 lm m  sr  at 555 nm ¼ 683 cd cm  at 555 nm
                  Radiant Intensity
                  1Wsr 21  ¼ 12.566 W (isotropic) ¼ 4p W ¼ 683 cd at 555 nm

                  Luminous Intensity
                  1lmsr ¼ 1cd ¼ 4p lm (isotropic) ¼ 1.464   10 23  Wsr 21  at 555 nm

                  Luminance

                  1lmm 22  sr 21  ¼ 1cdm 22  ¼ 10 24  lm cm 22  sr 21  ¼ 10 24  cd cm 22

                  Geometries
                  Converting between geometry-based measurement units is difficult, and should only be attempted
                  when it is impossible to measure in the actual desired units. You must be aware of what each of the
                  measurement geometries implicitly assumes before you can convert. The example below shows the
                  conversion between lux and lumens.
                     If you measure 22.0 lux from a light bulb at a distance of 3.162 m, how much light, in lumens, is
                  the bulb producing? Assume that the lamp is an isotropic point source, with the exception that the
                  base blocks a 308 solid angle. Using Equation (5.15), the irradiance at 1.0 m is E 1.0 m ¼ (3.162/
                     2
                  1.0)   22.0 ¼ 220 lm m 22 . On the basis of steradian definition [Equation (5.4)], we know that
                                                       2
                  1 sr cuts off a spherical surface area of 1 m at a distance of 1 m from the source. Therefore,
                         22                     21                                    21
                  220 lm m  corresponds to 220 lm sr  . The solid angle of the lamp is equal to 2p hr  [Equation
                  (5.4)], where h is the height of the spherical calotte, and corresponds to 2p   [1–cos (360 2 30/
                  2)] ¼ 12.35 sr, while the shadowed solid angle corresponds to 0.21 sr. Therefore, the total lumen
                  output will be 220 lm * sr 21 12.35 sr ¼ 2717 lm, (Figure 5.13).

                  PAR DETECTORS
                  Photosynthetic irradiance is the radiant flux density of PAR and is expressed as the radiant energy
                  (400–700 nm) incident on a unit of surface per unit time. A PAR detector is typically an irradiance
                  detector that is equally sensitive to light between 400 and 700 nm and insensitive to light outside
                  this region. Irradiance is now internationally expressed in moles of photons per unit area and
                  per unit time as mmol m 22  sec 21  (formerly mEinsten m 22  sec 21 ), where 1 mmol (mEinsten) corre-
                  sponds to 1 micromole of photons, that is, 6.02   10 17  photons, at a given wavelength. Modern
                  instruments measure in situ irradiance flux densities directly in mmol m 22  sec 21 .
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