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274                                   Algae: Anatomy, Biochemistry, and Biotechnology

                  then dried, either by spreading and air-drying or by using rotary dryers, and exported to the U.S. and
                  Denmark for processing.
                     Carrageenan production (valued at U.S. $240 million annually) was originally dependent on
                  wild macroalgae, especially Chondrus crispus (Irish moss), a small macroalgae growing in cold
                  waters, with a limited resource base in France, Ireland, Portugal, Spain, and the east coast provinces
                  of Canada. As the carrageenan industry expanded, the demand for raw material began to strain the
                  supply from natural resources. However, since the early 1970s the industry has expanded rapidly
                  following the availability of other carrageenan-containing macroalgae that have been successfully
                  cultivated in warmwater countries with low labor costs. Today, most raw material comes from two
                  species originally cultivated in the Philippines, Kappaphycus alvarezii and Eucheuma denticula-
                  tum, but which are now also cultivated in other warm-water countries such as Indonesia and the
                  United Republic of Tanzania. Limited quantities of wild Chondrus are still used; attempts to culti-
                  vate Chondrus in tanks have been successful biologically, but it has proved uneconomic as a raw
                  material for carrageenan. Wild species of Gigartina and Iridaea from Chile are also being harvested
                  and efforts are being made to find cultivation methods for these.
                     Table 7.6 summarizes commercially exploited algae and the corresponding extract.


                  ANIMAL FEED
                  Microalgae are utilized in aquaculture as live feeds for all growth stages of bivalve molluscs (e.g.,
                  oysters, scallops, clams, and mussels), for the larval/early juvenile stages of abalone, crustaceans,




                           TABLE 7.6
                           Summary of Commercially Exploited Algae and the Corresponding
                           Extracts
                           Scientific Name                Class                Extracts

                           Gracilaria chilensis       Floridophyceae       Agar
                           Ahnfeltia plicata          Floridophyceae       Agar/Carrageenan
                           Gelidium lingulatum        Floridophyceae       Agar
                           Pterocladia spp.           Floridophyceae       Agar
                           Hypnea spp.                Floridophyceae       Agar
                           Chondrus crispus           Floridophyceae       Carrageenan
                           Gigartina skottsbergii     Floridophyceae       Carrageenan
                           Gigartina canaliculata     Floridophyceae       Carrageenan
                           Mazzaella laminaroides     Floridophyceae       Carrageenan
                           Sarcothalia crispata       Floridophyceae       Carrageenan
                           Kappaphycus alvarezii      Floridophyceae       Carrageenan
                           Eucheuma denticulatum      Floridophyceae       Carrageenan
                           Iridaea spp.               Floridophyceae       Carrageenan
                           Laminaria hyperborea       Phaeophyceae         Alginate
                           Laminaria digitata         Phaeophyceae         Alginate
                           Laminaria japonica         Phaeophyceae         Alginate
                           Laminaria saccarina        Phaeophyceae         Alginate
                           Macrocystis pyrifera       Phaeophyceae         Alginate
                           Ascophyllum nodosum        Phaeophyceae         Alginate
                           Durvillea potatorum        Phaeophyceae         Alginate
                           Ecklonia spp.              Phaeophyceae         Alginate
                           Lessonia nigrescens        Phaeophyceae         Alginate
                           Lessonia trabiculata       Phaeophyceae         Alginate
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