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Chapter | 3  Biomass Characteristics                          55


             3.3.1 Structure of Wood
             Wood is typically made of hollow elongated and spindle-shaped cells
             arranged parallel to each other. Figure 3.4 is a photograph of the cross-
             section of a tree trunk showing the overall structure of a mature tree wood.
                Bark is the outermost layer of a tree trunk or branch. It comprises an
             outer dead portion and an inner live portion. The inner live layer carries food
             from the leaves to the growing parts of the tree. It is made up of another
             layer known as sapwood, which carries sap from the roots to the leaves.
             Beyond this layer lies the inactive heartwood. In any cut wood, we can easily
             note a large number of radial marks. These radial cells (wood rays) carry
             food across the wood layers.
                Wood cells that carry fluids are also known as fibers or tracheids.They are
             hollow and contain extractives and air. These cells vary in shape but are gener-
             ally short and pointed. The length of an average tracheid is about 1000 μmfor
             hardwood and typically 3000 8000 μm for softwood (Miller, 1999).
                Tracheids are narrow. For example, the average diameter of the tracheid of
             softwood is 33 μm. These cells are the main conduits for the movement of sap
             along the length of the tree trunk. They are mostly aligned longitudinally, but
             there are some radial tracheids that carry sap across layers. Lateral channels,
             called pith, transport water between adjacent cells across the cell layers.
             Softwood (coniferous) can have cells or channels for carrying resins. A hardwood
             (deciduous), on the other hand, contains large numbers of pores or open vessels.
                The tracheids or cells typically form an outer primary and an inner sec-
             ondary wall. A layer called the middle lamella joins or glues together
             the adjacent cells. The middle lamella is predominantly made of lignin.


                                                                Wood rays

                                                               Heartwood


                                                                Live bark




                                                                 Dead bark


                                                                Sapwood



             FIGURE 3.4 Cross-section of a tree trunk showing outer dead bark, inner live bark, sapwood,
             heartwood, and wood rays. Source: Photograph by author P. Basu.
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