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Figure 15.6 Hairy (a–d) and smooth (e–h) attachment systems. (a–b, e–f) Diagram of the action of both types of
systems on the smooth (a, e), and structured (b, f) substrata. (From Gorb, S.N. (2001) Attachment Devices of
Insect Cuticle. Dordrecht, Boston, London: Kluwer Academic Publishers. With permission of Springer Science þ
Business Media B.V.) Both systems are able to adapt to the surface profile. (c) SEM micrograph of the ventral
surface of the tarsus in the beetle Cantharis fusca (Coleoptera, Polyphaga). (d) SEM micrograph of the
ventral surface of the tarsus in the dobsonfly Sialis lutaria (Megaloptera). (g) SEM micrograph of the pretarsus of
the cicada Cercopis vulnerata (Auchenorrhyncha). (h) Sagittal section of the arolium of the locust Schistocerca
gregaria (Caelifera). AR, arolium; black arrow, ventral direction; HS, hairy soles; RD, filaments; TAR, tarsal
segments; UN, claw.

