Page 186 - Carbonate Facies in Geologic History
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Tubular Foraminifera, Komia, Tubiphytes                           173

                  2.  They  also  form  low-lying  plumose,  hemispherical  masses  interstratified  with  blue-
               green  algal  stromatolites  or  codiaceans.  The  algae  are  generally  coarse,  neomorphically
               crystallized, and most internal structure is lost. Such structure of "forams" resemble modern
               Nubicularia and also Renalcis from the older Paleozoic. Although the growth form is low and
               encrusting, as much as 8 m of such rock has been measured topping algal plate mud mounds.
               It forms  a true boundstone. Dunham (1969b)  has demonstrated  that it  and  Tubiphytes  are
               responsible for growth constructed cavities in the Townsend-Kemnitz reservoir in Lea Coun-
               ty, New  Mexico. When weathered,  the  coarser crystallized  algae take  on  a  clear, white,  or
               yellow  (iron-stained) color and the foraminifera are black.  The foraminifera  have  micropo-
               rous tests and, when in unweathered rock, are white. This characteristic rock caps bioherms
               on the seaward or west side of the Sacramento Virgilian complex. It also forms  small down-
               slope bioherms, "satellites," on slopes within flank beds of larger mounds. Recent opthalmids
               are known in Florida waters but are not abundant (Moore, 1957). They flourish only 7-10 m
               deep in the relatively quiet shelf water of the Florida Reef Tract, behind the reefs  and down
               the foreslope of the reef to 40 m depth. They occur in water  of normal  marine salinity and
               circulation. Bioherms in the outcropping Paradox Formation of Utah are composed chiefly
               of such forms intergrown with plumose algae (codiaceans?) and contain abundant scattered
               Chaetetes and Caninia.



               Tubiphytes  Maslov,  1956  (Nigriporella  of Rigby,  1958;  and  Hydrocorallines  of
               Newell et aI., 1953) (Plates IV A, XXIV B)

               The  form  is  defined  as  tiny  laminated  growths  which  are,  like  the  tubular  foraminifera,
               epiphytic,  encrusting and  of low  relief.  Their  biological  affinity  is  not  known.  They  were
               originally described in Russia (Shamovella, Rauser-Chernoussova, 1951  n.nudem and as  Tubi-
               phytes  Maslov,  1956)  and  later  termed  Nigriporella  by  Rigby  who  believed  them  to  be
               perhaps  calcified  tunicates.  A  good  review  of  the  genus  was  published  by  Toomey  and
               Croneis, 1965, who consider them algae. They commonly contain clear calcite areas about
               which faint laminae are concentric. These areas may be openings within the structure of the
               organism (conceptacles?)  or  merely  holes,  representing vanished  elements  of the encrusted
               biota. Several holes may be seen within a single laminar mass. Like the tubular foraminifera,
               Tubiphytes  was microporous (porcellaneous). It is  dead white in  reflected  light  and dark in
               transmitted light. Silicified material from the Big Hatchet Mountains, New Mexico, indicates
               that the organism  may  occur  in  small  irregular  balls  a  centimeter  or less  in  diameter and
               arranged in chains or baskets.
                  On a world-wide basis, Tubiphytes occurs first in the early Carboniferous and is common
               into Late Jurassic reefoid strata. The type is of Kungurian (Permian) age. In the southwestern
               U.SA it appears in the Late Pennsylvanian, but becomes an important rock builder only in
               the Wolfcampian where  it may dominate the biota  of bioherms, growing  intertwined with
               tubular foraminifera  and  existing  in  flank  beds  whose  sediments  derived  from  organisms
               growing on top of the mounds. The exact ecological position of Tubiphytes within such reefs is
               not certain; it is believed to form a boundstone with tubular foraminifera on exposed crests
               and seaward flanks of many carbonate buildups.



               Komia Korde, 1951, a Tiny Dendroid Stromatoporoid (Plate XXIII B)

               Wilson et aI., (1963) gives a description and proper taxonomy. Komia was assigned to the red
               algae by Johnson (1961). Toomey and Johnson erroneously assigned certain branched forms
               of Komia  to  Ungdarella  (Wilson,  1969).  It apparently has  a  branched  and flabellate  form.
               Mostly it exists as tiny broken twigs much like the larger Devonian Stachyoides  and Amphi-
               pora which it greatly resembles in internal  structure.  Komia  is  generally  present  in  bedded
               limestones in the Middle Pennsylvanian (Derryan and Des Moinesian) in the southwest. In
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