Page 254 - Carbonate Facies in Geologic History
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Organic Composition of Ladinian Wetterstein Limestone             241

                  The Wetterstein limestone forms  major offshore banks like those of the Do-
               lomites but is much less dolomitized and has been carefully studied petrographi-
               cally in recent years (Ott, 1967; Sarntheim, 1967; Toschek, 1968). The limestone is
               hundreds  of meters  thick  (up  to  1500 m)  and  shows  generally  regular  upward
               shoaling major facies  which also may be traced laterally in  belts  surrounding a
               series of east-west elongate carbonate banks (Fig. VIII-16). The exact distribution
               of the  several  known  banks  is  debated  because  of  tectonic  displacements  so
               common  in  this  Alpine  structure belt.  The  Karwendel  bank,  forming  the  high
               cliffs north of Innsbruck, is best known. Its boundary is a bioclastic girdle, wider
               on the south and perhaps indicating stronger wind waves and currents from that
               direction. Its narrow northern boundary is also clearly defined.  Other banks are
               recognized south of Innsbruck.
                  All  of these  banks  rise  from  a  platform  limestone  of Anisian  age-termed
               Alpine Muschelkalk. Their equivalent basinal facies (Partnach beds) is  only  100-
               250 m thick and is dark, thin-bedded, slightly bioclastic-spiculitic limestone with
               chert nodules and several shale beds. This grades upward into the  bank proper
               through some  thick-bedded, dark gray,  bioclastic wackestone with  peloids  and
               abundant crinoid stems, mollusks, and dasycladaceans derived from upslope. No
               real forereefbreccia is present (unlike the beds in the Dolomites). The Wetter stein
               transition beds instead are a type of basin marginal deeper water limestone (Fa-
               cies  belt 3  of the  author).  Texturally,  the  bank  edges  consist  of  poorly  sorted
               coarse  bioclastic  debris  generally  in  a  sparry  matrix  but  with  some  bioclastic
               micritic infill. Much coarse spar coating is seen. Texture is essentially rudstone to
               floatstone. The biota of this bank edge facies  includes large  crinoids, echinoids,
               Solenopora,  Daonella  bivalves,  the  brachiopod  Rhynchonella,  gastropod  Euom-
               phalus, orthoconic nautiloids, dasycladacean Teutloporella nodosa, some few  cor-
               als, many segmented or beaded  sponges,  and encrusting  Tubiphytes  (Plates  XX,
               IV B, XXV A, XXVI A).
                  Ott's  careful  petrographic  study  (1967)  shows  that  Tubiphytes  is  the  most
               important reef builder. He lists five  different finely  textured, encrusting growths
               with delicate, tiny (2 mm) bushy networks in the sedimentary fabric. Microscopic
               study also discerns minute,  1 mm thick, stalks of crystalline  calcite  and includes
               tiny porous pipes in fasciculate or isolated form  up to 3 mm in diameter. Those
               with nonporous walls are termed H olocoelia. Very delicate fine-branching septate
               corals, Calamophyllia, are up to 3 mm in  diameter and encrust the epithecae  of
               other forms. Larger forms are also present. Irregular meshworks and thick stalks
               of H olocoelia are present as  are fasciculate forms  such as "1hecosmilia", M arga-
               rosmilia,  and Pinacophyllum.  The  relative  abundance  of  such  corals  is  hard  to
               estimate. The megafauna is dominated by diverse groups of calcareous sponges.
               Inozoa or pharetrones are common but eight genera of columnar, segmented or
               beaded  forms  of Sphinctozoans  have  been  described  (Ott,  1967).  The  sponges
               seem to have grown in groups comprised  of only a limited number of genera at
               one place and to have occupied different ecologic niches from the corals. Hydro-
               zoans, both as tiny stalks and as encrusting concentric balls, are present but not
               common. Some few encrusting calcareous foraminifera and bryozoans appear to
               have  been  sponge  parasites.  Calcareous  algae  are  not  particularly  important;
               some blue-green stromatolitic forms  may  occur  as  reef binders.  The  codiacean
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