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The Evolution of Cultures  435

            Most of these ancestral bands probably also engaged in skirmishes
        along territorial boundaries initiated by groups of males. Physical strength,
        resilience to disease, and capacity for coordinated collaborative action were
        factors that jointly determined the success in these actions. Larger bands
        would have assimilated smaller ones, sometimes by killing adult males and
        infants and adopting females. Successful groups would have split up when
        an aspiring leader left the mother group, followed by part of the group
        members. 6
            So, fi ssion and fusion of groups has been a mechanism for combining
        genetic and cultural evolution for millions of years. The normal course of
        evolution among apes has been that separation of subpopulations gradu-
        ally led to the formation of new races and, after millions of years, new
        species—a process called speciation. In contrast, the splitting and merging
        and the exchange of females among our ancestors would have tended to
        promote genetic similarity but cultural variation between groups, instead
        of genetic divergence and speciation as is usual in the animal kingdom.
        Fossil evidence is sketchy and still growing, which means that naming is
        still controversial and subject to revision. So far, the evidence suggests that
        a few million years ago, several isolated populations of hominins existed
        in what is now Africa. They created diversity in genetic materials and
        developed into very different forms—for example, the recently announced
        “Ardi” (Ardipithecus ramidus), who lived 4.4 million years ago in what is
        now Ethiopia, or the well known “Lucy” (Australopithecus afarensis), who
        lived in the same area a million years later. Many lineages have died out
        since then. No traces of anything but the crudest tool use were found.
            From around 1.8 million years ago, brain size started to grow rapidly.
        This development was probably associated with refinements in communi-

        cation skills and in what evolutionary psychologists call “theory of mind.”
        Theory of mind is the level of understanding of the beliefs, desires, and

        intentions of others. Today’s humans can understand utterances fi ve levels
        of mind deep—for example, “I believe that you think that he hoped that
        she would rejoice at his distress.” Theory of mind is crucial for maintaining
        moral circles in a socially complex world, and it has gradually developed
        during our history.
            There are still many guesses concerning this period, such as about
        how soon and how often hominins left Africa. The forebears of Homo erec-
        tus may have left Africa 1.8 million years ago, giving rise over several hun-
        dred thousand years to Homo heidelbergensis, which later evolved into Homo
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