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CHAPTER 15 • Humans and Preindustrial Climate  277


                                         Ice volume         America, and North America. This change altered the
                                            18
                Dust                       (δ O)            carbon-isotopic composition of the carbon left behind in
            More        Less           Max.           Min.
           0                                                soils and in the teeth of grazing animals on all four conti-
                                                                                          13
                                                            nents. The trend toward heavier δ C values in North
                                                            Africa (Figure 15–5 center) indicates a change from C3
                                                            vegetation (trees and shrubs) toward C4 vegetation
                                                            (warm-season grasses). Greenhouse experiments suggest
                                                            that such a shift could have happened because CO con-
           1                                                                                          2
                                                            centrations fell past a threshold of 600–500 ppm over
                                                            several million years. The fact that a gradual change from
                                                            C3 to C4 carbon occurred on all four continents suggests
                                                            falling atmospheric CO concentrations as a common
                                                                                2
                                                            explanation. As this trend developed, global climate was
           2                                                cooling, and ice sheets were growing (Figure 15–5 right).
                                                               Climate modeling points to several factors that may
          Myr ago                                           have driven a long-term drying in Africa. One factor
                                                            was construction of volcanic plateaus in east Africa over
                                                            the last 30 million years. Model simulations show that
                                                            prior to plateau construction, moisture-bearing winds
           3
                                                            blew from the Indian Ocean westward into East Africa
                                                            (Figure 15–6). As the plateaus took form, this influx of
                                                            moisture was choked off, and the vegetation shifted
                                                            from forests to grasslands across much of the region
                                                            where most hominin remains are found but not across
           4                                                the rest of Africa. Another factor that may have con-
                              Carbon
                       Grassland    Woodland                tributed to this regional drying was a cooling of the


        FIGURE 15-5 Long-term changes in African dust and
        vegetation Over the last 4.5 Myr, (left) increasing amounts of
        dust were blown from North Africa to the tropical Atlantic,                  Inland sea
        and (center) vegetation cover in many regions gradually shifted
        away from trees and shrubs toward warm-season grasses.
        (Right) These changes occurred during a time of global
        cooling. (Adapted from P. B. deMenocal, “Plio-Pleistocene
        African Climate,” Science 270 [1995]: 53–59.)                                            Tibet





        also have increased because the spring winds that lift
        and carry the fine debris became stronger.
           A long-term drying trend in North Africa is also
        evident in a sediment sequence from the Atlantic Ocean
        just offshore of the Sahara Desert that contains a his-
                                                            FIGURE 15-6 Tectonic effects on African climate Over
        tory of pollen changes over portions of the last 3.7 Myr.
                                                            the last 20 Myr, East African plateaus became drier because of
        Gradually decreasing amounts of forest pollen and
                                                            local uplift in eastern Africa and cooling of the western Indian
        increasing amounts of savanna and desert scrub pollen
                                                            Ocean (orange), uplift of the Tibetan Plateau (yellow), and
        indicate a progressive trend toward dry-adapted vegeta-
                                                            shrinkage of an inland sea in west-central Asia (blue).
        tion. Other shorter-term pollen records from the East  (Adapted from P. Sepulchre et al., “East African
        African plateau also reveal a long-term trend toward  Aridification and Late Neogene Uplift,” Science 313 [2007]:
        more open vegetation.                               1419–23, W. F. Ruddiman and J. E. Kutzbach, “Plateau Uplift
           A long-term decrease in atmospheric CO could also  and Climate Change,” Scientific American 264 [1991]: 66–75,
                                              2
        have been a factor in the change in vegetation. Over this  and G. Ramstein et al., “Effect of Orogeny, Plate Motion, and
        interval, a major vegetation shift was underway not just in  Land-Sea Distribution on Eurasian Climate over the Past 30
        East Africa but also in parts of southern Asia, South  Million Years,” Nature 386 [1997]: 786–95.)
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