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278     PART V • Historical and Future Climate Change


        western Indian Ocean caused by the partial closure of a  open vegetation at or near the onset of the glacial cycles.
        seaway near Indonesia and the resulting loss of warm  Subsequently, forest vegetation repeatedly fluctuated up
        tropical water from the western Pacific Ocean.      and down the sides of the mountains at intervals of tens
           Tectonic changes in southern Asia may have also  of thousands of years. These oscillations appear to be
        played a role. Continuing uplift of the northern and  connected to the early northern hemisphere glacial
        eastern parts of the Tibetan Plateau not only strength-  cycles.
        ened the Asian summer monsoon but also created a       In addition, records from the eastern and southern
        strong counterclockwise spiral of winds that drove hot,  plateaus of Africa also show a widespread change from
        dry air out of the interior of Asia across Arabia and into  woodland-adapted browsing (leaf-eating or grass-eating)
        northeastern Africa (Figure 15–6). These winds pro-  animals to grazing animals strictly adapted to grasslands
        duced much drier conditions in summer across a huge  near and after 2.5 Myr ago. Many browsing animals
        arc extending from west-central Asia across the Arabian  became extinct, and new grazing animals appeared both
        peninsula and into northeastern Africa. This drying  by evolving and by emigrating from Asia. The evidence
        trend may have been aided by the retreat of a vast inte-  suggests a relatively rapid proliferation of open grass-
        rior seaway that once occupied west-central Asia and  lands at the expense of closed forests. This marked
        moderated its temperatures.                         change from browsers to grazers (and from forest to
                                                            grassland habitat) has been attributed to the cooling and
          IN SUMMARY, both the available data and model     drying produced by northern hemisphere glacial cycles.
          simulations confirms a gradual trend toward more     One problem with this proposed link to the north-
          open vegetation in Africa throughout the time of  ern ice sheets is that GCM simulations suggest that the
          human evolution. This evidence seems to support   ice sheets would have had relatively small direct effects
          the savanna hypothesis.                           on temperature and precipitation in North Africa. Even
                                                            the large ice sheets that existed at the last glacial maxi-
           The actual record of hominin remains, however,   mum, 20,000 years ago, cooled northern Africa by no
        suggests that the picture is more complicated. Hominins  more than 1 –4 C, with smaller changes farther south
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        occupied environments ranging from woodlands and    and little change anywhere during the summer (mon-
        grasslands to river margins throughout their history,  soon) season. The smaller ice sheets that grew and
        with no obvious trend toward greater occupation of  melted during the glacial cycles that started 2.75 Myr
        the savanna environment. This evidence indicates that  ago would probably have had even smaller impacts on
        hominins have long been resourceful in making use   African climate.
        of a range of environments and it has led to a different  It is possible that these early glaciations had some
        hypothesis of the effects of climate on human evolution.  effect on vegetation and habitat in Africa because they
           The basic premise of the  variability selection  were accompanied by drops in atmospheric CO levels,
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        hypothesis is that rapid evolution occurred because  like those that have occurred during the last several
        alteration of habitat put new demands on our ancestors,  hundred thousand years (see Chapter 10). Lower CO
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        some of whom then took advantage of new opportuni-  levels during these earlier glaciations could have caused
        ties. Changes in climate favored hominims with traits  some replacement of C3 trees and shrubs by C4 grasses
        useful for survival in new environments, particularly  during each glacial cycle, with fragmentation of forest
        clans or tribes that had larger numbers of members with  habitat.
        such traits.                                           Another problem with the variability selection
           Climatic variability has greatly increased at high lat-  hypothesis is that the summer monsoons that control
        itudes during the last several million years, particularly  annual rainfall and vegetation types across the savannas
        with the onset of small glaciation cycles near 2.75 Myr  south of the Sahara Desert and across most of the
        ago and then the larger oscillations since 0.9 Myr ago  plateaus of East Africa appear to have continued for
        (Chapter 9). In addition, the earliest appearance of our  many millions of years with no strong trend toward
        genus Homo is dated to between 2.4 and 2.0 Myr ago,  larger or smaller oscillations. It appears that the mon-
        not long after the first glacial cycles. Perhaps these early  soons did not contribute to a long-term increase in
        glaciations created cycles of cooling and drying in  climatic variability.
        Africa that accelerated the pace of evolution by favoring  Unfortunately, the fossil record of human remains is
        individuals with greater adaptability.              too sparse to determine any correlation between the
           Several lines of evidence suggest that northern  onset of glacial cycles and hominid evolution.
        hemisphere glaciation could have affected climate and
        hominin habitat in Africa. Pollen records from high ter-  IN SUMMARY, the record of human remains is too
        rain in East Africa show that the upper tree line began  sparse to test the variability selection hypothesis.
        to retreat up mountainsides and give way to more
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