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64   CHAPTER 3



                                                        stronger competition. Conversely, continental rifting
                                                        leads to the isolation of faunas which then follow
                                                        their own distinct evolutionary development. For
                                                        example, marsupial mammals probably reached Aus-
                                                        tralia from South America in the Upper Cretaceous
                                                        along an Antarctic migration route (Hallam, 1981)
                                                        before the Late Cretaceous marine transgression
                                                        removed the land connection between South America
                                                        and Antarctica and closed the route for the later
                                                        evolving placental mammals. Sea floor spreading then

                                                        ensured the isolation of Australia when the sea level
                                                        dropped, and the marsupials evolved unchallenged
                                                        until the Neogene when the collision of Asia and
                                                        New Guinea allowed the colonization of placental
           Figure 3.11  Correlation of invertebrate diversity with   mammals from Asia.
           time and continental distribution. A, earlier Pangea;
           B, fragmentation of earlier Pangea producing oceans
           preceding Caledonian (1), Appalachian (2), Variscan (3),
           and Uralian (4) orogenies; C, suturing during Caledonian   3.6 PALEOMAGNETISM
           and Acadian orogenies; D, suturing during Appalachian
           and Variscan orogenies; E, suturing of Urals and
           reassembly of Pangea; F, opening of Tethys Ocean;   3.6.1 Introduction
           G, fragmentation of Pangea. a, Gondwana; b, Laurasia;
           c, North America; d, South America; e, Eurasia; f, Africa;
           g, Antarctica; h, India; i, Australia (after Valentine &   The science of paleomagnetism is concerned with
           Moores, 1970, with permission from Nature 228, 657–9.   studies of the fossil magnetism that is retained in certain
           Copyright 1970 Macmillan Publishers Ltd).    rocks. If this magnetism originated at the time the rock
                                                        was formed, measurement of its direction can be used
                                                        to determine the latitude at which the rock was created.
                                                        If this latitude differs from the present latitude at which
           species as a result of genetic isolation and the mor-  the rock is found, very strong evidence has been fur-
           phological divergence of separate faunas. Conse-  nished that it has moved over the surface of the Earth.
           quently, more species evolve as different types occupy   Moreover, if it can be shown that the pattern of move-
           similar ecological niches. Figure 3.11, from Valentine   ment differs from that of rocks of the same age on a
           & Moores (1970), compares the variation in the   different continent, relative movement must have
           number of fossil invertebrate families existing in the   occurred between them. In this way, paleomagnetic
           Phanerozoic with the degree of continental fragmen-  measurements demonstrated that continental drift has
           tation as represented by topological models. The cor-  taken place, and provided the first quantitative estimates

           relation between number of species and fragmentation   of relative continental movements. For fuller accounts
           is readily apparent. An example of such divergence   of the paleomagnetic method, see Tarling (1983) and
           is the evolution of anteating mammals. As the result   McElhinny & McFadden (2000).
           of evolutionary divergence this specialized mode of
           behavior is followed by different orders on separated
           continents: the antbears (Edentata) of South America,
           the pangolins (Pholidota) of northeast Africa and   3.6.2 Rock magnetism
           southeast Asia, the aardvarks (Tubulidentata) of central
           and southern Africa, and the spiny anteaters (Mono-  Paleomagnetic techniques make use of the phenome-
           tremata) of Australia.                       non that certain minerals are capable of retaining a
             Continental suturing leads to the homogenization   record of the past direction of the Earth’s magnetic

           of faunas by cross-migration (Hallam, 1972) and the   field. These minerals are all paramagnetic, that is, they
           extinction of any less well-adapted groups which face   contain atoms which possess an odd number of elec-
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