Page 180 - Introduction to Paleobiology and The Fossil Record
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MASS EXTINCTIONS AND BIODIVERSITY LOSS  167


                                     Sediment type


                                Zone
                                C28R       Sample numbers  Brazos core,
                                                    Texas
                           Lower Tertiary (Danian) P1a                                      P1b  C29N











                                C29R 1.0                       0.5                       0                        0.5                      1.0                     1.5m 217                       225                     242             256                  268        279                                289  KT boundary
                              P0



                           Upper Maastrichtian  Pseudotextularia deformis








                                C30N



             Figure 7.4  Patterns of extinction of foraminifera in a classic KT section spanning about 1.5 myr. A
             species loss of 53% occurred in two steps close to the KT boundary and iridium anomaly. Dating is
             based on magnetostratigraphy, and the KT boundary falls in the C29R (reversed) zone. Planktonic
             zones (P0, P1a, P1b) are indicated; sediment types are mudstones (darker grey) and limestones (pale
             grey); meter scale bar shows height above and below a particular extinction level, 0. (Based on Keller
             et al. 1993.)

             answers to questions such as these refer to     raphy (see p. 24) and exact radiometric dating
             ecological time scales – that is, times of years   (see p. 38) are also needed.
             or decades at most.

               It is just as difficult, if not more so, to
             answer questions of the timing of ancient
             events from region to region or continent to    Selectivity and mass extinctions
             continent. How can a paleontologist be sure     The second defi ning character of mass extinc-
             that the supposed KT boundary in Montana        tions (see p. 164) was that they should be
             is the same as the supposed KT boundary in      ecologically catholic, that there should be
             Mongolia? Perhaps the boundary is marked        little evidence of selectivity. Ecological selec-
             as the next sedimentary rock layer above the    tivity implies that some organisms might be
             appearance of the last dinosaur fossil. But of   better able to survive a mass extinction event
             course this definition is perfectly circular: the   than others. Mass extinctions do not seem to

             KT boundary is marked by the disappearance      have been particularly selective, even though
             of dinosaurs; dinosaurs disappeared just        it might seem that, for example, large reptiles
             below the KT boundary. Other fossils, such      were specially selected for extinction during
             as pollen, may be used to date the boundary,    the KT event. The dinosaurs and some other
             but additional evidence, from magnetostratig-   large reptiles certainly died out then, but a
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