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376  INTRODUCTION TO PALEOBIOLOGY AND THE FOSSIL RECORD


                                              Number of        Number of            Vanadium/(Vanadium+Nickel)
                                            olenid specimens  agnostid specimens             ratio
                                           0 5 10 15 20 25 30 35  0 5 10 15 20 25 30 35 40  (m)0.85  0.80  0.75  0.70
                                        (cm)
                                        230
                                 Material collected by Lauridsen and Nielsen (2005)  O. dentatus  200  Upper O. attenuatus  Faulted cut in drill core  (m)
                                        220

                                        210


                                        190
                                        180
                                        170
                                        160
                                                                                                      –78.0
                                      O. attenuatus
                                        150
                                        140
                                                                                                      –78.2
                                        130
                                        120       Cyclotron sp.           Lower O. attenuatus         –78.1
                                                                          No agnostic or              –78.3
                                                                          Olenus species
                                        110
                             Material collected by Clarkson et al. (1998)  O. transversus – O. truncatus – O. wahlenbergi  90  Range of O. truncatus  –78.5
                                                                                                      –78.4
                                        100
                                                                          Sole presence of
                                                                               agnostids
                                         80
                                         70
                                                                                                      –78.6
                                         60
                                                                                                      –78.7
                                         50
                                                                          to O. wahlenbergi
                                                                                                      –78.8
                                         40
                                         30
                                         20
                                                                                                      –79.0
                                         10                                                           –78.9
                                                                          O. transversus
                                          0
                            Limestone                                                                 –79.1
                             interval                                     Limestone
                           not sampled  O. gibbosus                                                   –79.2
                              Material collected  by Lauridsen and  Nielsen (2005)  –30  Range of O. gibbosus  –79.3

                                        –40
                                                                                                      –79.4
                                        –50
                                            Complete specimen    Cephalon, Homagnostus
                                            Cranidium, Olenus    Pygidium, Homagnostus
                                            Pygidium, Olenus     Cephalon, Glyptagnostus
                                            Loose cheek, Olenus  Cyclotron sp. (bradoriid)
                                            No data at this level

                      Figure 14.13  Microevolution and faunal dynamics of olenids in the Swedish Alum Shales. Olenus
                      species evolve gradually up through the section. (Based on Clarkson et al. 1998.)
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