Page 235 - Petroleum Geology
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            ment deficient in oxygen, and that this organic matter was derived from the
            tissues of living plants and animals.
              When living tissues die, they are either consumed  by living organisms  and
            pass into the food chain, or they  decompose with the aid of bacteria. If the
            environment  is  sufficiently  rich  in  oxygen,  those  tissues not consumed by
            scavengers decompose to CO,  and water (mainly): if  the environment is defi-
            cient  in  oxygen  and free of scavengers, the products are more complicated
            and varied, but are essentially compounds of carbon, hydrogen, oxygen, and
            nitrogen.
              We  concluded  in  the general review that the lack of  correspondence be-
            tween the composition  of  crude oils and the nature of  organic life on earth
            when the crude oil was presumably formed, revealed by the fossils, indicated
            a source for hydrocarbons in the fundamental biological molecules: proteins
            (amino acids), lipids (fats, vegetable oils, waxes), carbohydrates, and, in the
            higher plants,  lignins. Some of these are present in all forms of life, and their
            initial preservation, which is necessary for their subsequent alteration, depends
            on  their rapid  transport  to an environment  free of  scavengers and deficient
            in oxygen, and their incorporation there into the accumulating sediment. This
            anoxic  environment  is  not  necessarily  at the  sea water/sediment  interface,
            but it must exist at a very shallow depth of burial. What matters is the balance
            between the amount of material to be oxidized and the available oxygen, be-
            cause a  large  supply of  organic material can create its own anoxic environ-
            ment.  These environments will be free from scavenging animals, but not free
            from bacteria.
              The physical environment most prone to develop anoxic chemical environ-
            ments is one of little energy, that is, one in which only fine-grained sediment
            is present. Thereafter, the preservation of organic matter in quantity depends
            on its being buried as it is supplied, that is, the area must be one accumulating
            fine-grained sediment in a sedimentary  basin. Note that the accumulation of
            fine-grained sediment is not in itself sufficient: there must also be a supply of
            organic  matter to an anoxic environment.  So true source rock must be  dis-
            tinguished  from source rock  in a broad sense because not all of a mudstone
            unit, for example, will necessarily be true petroleum source rock.
              The requirement of large supplies of organic matter over considerable peri-
            ods of  time implies that we are concerned with sedimentary basins near con-
            tinental margins mainly  (but not exclusively) because these are areas of large
            biomass  where  sediment  is  commonly  accumulating  into  the stratigraphic
            record.  In such areas, great variations  may exist between the relative supply
            of  terrestrial  and  marine  organic  matter, and these differ in their chemical
            compositions. For large supplies of  organic matter, organisms such as plank-
            tonic algae in the marine environment and higher plants in the terrestrial are
            more  likely  progenitors  than  vertebrates,  for  example.  Marine  planktonic
            algae contain variable amounts of proteins, lipids and carbohydrates, whereas
            higher  land  plants  contain  cellulose and  lignin,  with  some lipids in spores,
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