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PALEOECOLOGY AND PALEOCLIMATES  105




                                      Suspension  Herbivore   Carnivore    Modern
                                                                         evolutionary
                                                            Osteichthyes    fauna
                                     Malacostraca  Osteichthyes  Chondrichthyes
                                   Pelagic  Gastropoda  Mammalia  Mammalia
                                                             Reptilia
                                      Mammalia
                                                            Cephalopoda
                                     Suspension  Deposit     Herbivore  Carnivore
                                                            Gastropoda  Cephalopoda
                                                           Polyplacophora  Malacostraca
                                      Bivalvia  Gastropoda
                             Mobile                       Monoplacophora Echinoidea
                                      Crinoidea  Monoplacophora
                                                             Ostracoda  Stelleroidea
                                                            Echinoidea  Cephalopoda
                                      Bivalvia
                                      Articulata
                                      Anthozoa
                            Attached  Cirripedia                Suspension  Deposit  Carnivore
                              low
                                    Gymnolaemata                  Bivalvia
                                                         Passive
                                     Stenolaemata        shallow  Echinoidea  Bivalvia  Bivalvia
                         Epifauna   Gymnolaemata                 Gastropoda  Bivalvia  Gastropoda
                                      Polychaeta
                                                                  Bivalvia
                                                                           Echinoidea
                                                                 Polychaeta
                                                         active
                            Attached  Stenolaemata       Shallow  Echinoidea Holothuroidea  Malacostraca
                                      Anthozoa
                                                                                     Polychaeta
                             erect  Hexactineluda
                                    Demospongia                            Polychaeta
                                      Calcarea       Infauna  Deep
                                     Gastropoda          passive  Bivalvia
                            Reclining  Bivalvia
                                     Stelleroidea                 Bivalvia
                                      Anthozoa           Deep    Polychaeta  Bivalvia  Polychaeta
                                                         active            Polychaeta
                                                                 Malacostraca
                        (c)
                          100
                                 soft-part faunas
                                 skeletal faunas
                          80
                                 mean α
                         Number of modes of life  60    Phanerozoic trends
                          40
                          20

                           0
                            Early Late  E.M.  Late            Recent
                            Ediacaran Cambrian Ordovician     Neogene
                                         Geological time
                        (d)
               Figure 4.21  Continued





             During the last 600 million years the Earth     zones: humid tropical (no winters and average

             has oscillated at least five times between ice-  temperatures above 18°C), dry subtropical
             house and greenhouse conditions, spending       (evaporation exceeds precipitation), warm
             most of the time in greenhouse climates (Fig.   temperate (mild winters), cool temperate
             4.23). For much of the Precambrian the Earth    (severe winters) and polar (no summers and
             probably endured relatively cool climates.      temperatures below 10°C). But can these
             The Earth is generally divided into fi ve climate   zones be recognized through deep time and be
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