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292  INTRODUCTION TO PALEOBIOLOGY AND THE FOSSIL RECORD



                                                                                     small favositid and
                                                                                   heliolitid tabulate corals
                               clusters of crinoids



                                                                                              small stromatoporoids




                           generalist brachiopods
                              and molluscs


                                                                     fasciculate rugose corals
                           (a)
                                                    large, platy to domed favositid and
                                                       heliolitid tabulate corals
                                                                                             diverse crinoids

                            cerioid, astraeoid,
                            thamnasterioid and
                           aphroid rugose corals                                          massive stromatoporoids










                             specialist, small to large brachiopods
                                    and mollusks




                           (b)
                      Figure 11.34  Pioneer (a) and climax (b) reef communities in Silurian and Devonan reef systems. (From
                      Copper, P. 1988. Palaios 3.)

                      long and up to 300 km wide and visible from     facies can be mapped in considerable detail,
                      space. It is a long-lived structure dating back   as the Windjana Gorge dissects the near-
                      to the Miocene. The reef extends from 9˚ to     horizontal strata of the northern margin of
                      25˚ south and comprises many multicolored       the Canning Basin (Fig. 11.35). An unbedded
                      scleractinian corals together with many other   core of calcareous algae, corals and stromato-
                      invertebrates and calcareous algae. The fore-   poroids sheltered a back-reef and lagoonal
                      reef deposits tumble eastwards into the         environment packed with calcareous algae,

                      western Pacific; landward back-reef lagoons      corals, stromatoporoids and crinoids together
                      are developed against eastern Australia. Can    with brachiopods, bivalves, cephalopods and
                      such reef constellations really be recognized   gastropods. In front the fore-reef was steep
                      in the fossil record? On the adjacent continent   and littered by reef talus. However, during the
                      the Upper Devonian rocks of the Canning         Late Devonian extinction event, at the end
                      Basin contain fossil barrier reefs dominated    of the Frasnian, associations dominated by
                      by calcareous algae and tabulate and rugose     rugose and tabulate corals together with stro-
                      corals together with stromatoporoids and        matoporoids disappeared; this type of reef
                      microbiolites. The reef and its associated      ecosystem never recovered.
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