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            the remediation of degraded areas should be preceded by analyses in the hydroponic
            system and then field trials, in order to determine actual capacity to absorb toxic
            elements and to identify the effect of environmental conditions on plant behaviour
            (Zabłudowska et al. 2009). Another essential criterion is associated with biomass,
            since plants exhibiting high efficiency of heavy metal uptake are generally
            characterised by a slight increase in biomass (Herna ´ndez-Allica et al. 2008). In
            this respect full understanding of the genetic regulation of plant biomass production
            is crucial (Demura and Ye 2010). Within the last 20 years many different tools have
            been used to increase the efficiency of phytoremediation thanks to the application
            of specific additives in the form of complexing substances (Yan et al. 2012) and the
            application of bacteria.



            5.2  Role of Soil Conditions in Natural Plant Growth
                 (Macro- and Microelements)

            Soil is a very important natural element of the environment preconditioning plant
            life. It covers the earth crust (lithosphere) with a thin stratum 1.5–2.0 m thick which
            developed from definite parent material under the influence of soil-forming factors
            and processes. The essence of these processes consists in transformations of
            mineral (degradation and synthesis) and organic (mineralisation and humification)
            materials, horizontal and vertical dislocation of many chemical elements and
            compounds, as well as exchange of matter and flow of energy between live
            organisms and soil substrate or soil phases (solid, liquid, and gaseous)
            (Prusinkiewicz 1999). The coloured effect of soil-forming processes involves
            appropriate formation and arrangement of genetic horizons in the soil perpendicular
            cross section. The upper horizons, called epipedons, as well as internal horizons
            referred to as endopedons, are characterised by strictly defined parameters and exert
            a strong impact on the membership of a given soil in a specific taxonomic unit
            (FAO 2007; PTG 2011).
              From the point of view of plant growth and development as well as biomass
            production, near-surface horizons—frequently referred to as accumulative-humus
            horizons—play a particularly important role. They are made up of the following
            four components: mineral material and organic matter (solid phase), soil water
            (liquid phase), and air (gaseous phase) (Fig. 5.1). In mature soils, this triple-phase
            system remains in a certain state of dynamic equilibrium determining physical,
            chemical, and biological soil properties. A disturbance of this balance under the
            influence of natural factors is described as soil evolution, whereas any deterioration
            of soil properties as a result of anthropogenisation (human pressure) results in soil
            physical, chemical, or biological degradation. The most important constituents of
            the solid phase comprise mineral and organic colloids, i.e. particles of dimensions
            below 2 μM. Soils inherit their mineral colloid content together with their parent
            material from which they were formed. In the majority of cases, these are classic-
            sedimentary rocks and partially non-classic rocks, primarily, of post-glacial origin
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