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Modeling and control in physiology  25


              Table 3 Some basic sensors and effectors in the human body.
              Human        Body organs/
              transducers  tissues       Senses/aptitude/action

              Sensors      Eyes          Vision
                           Nose          Smell
                           Tongue        Taste
                           Ears          Hearing/static and dynamic balance
                           Skin          Touch, vibroception, thermoception,
                                           nociception, proprioception...
              Effectors    Skin and      Physical pressure and force
                             muscles
                           Sweat glands  Produce sweat (increases heat loss)
                           Pancreas      Produces insulin (regulates glucose in blood)
                           Skeletal      Guarantees postural balance
                             system

























              Fig. 20 Postural balance via sensors and effectors; (A) schematic model (B) block
              diagram. (The figure has been adopted from Kim, S., et al., 2009. Postural feedback
              scaling deficits in Parkinson’s disease. J. Neurophysiol. 102 (5), 2910–2920.)



              sensors to measure and anticipate external perturbations as is shown in Fig. 21.
              An example for voluntary movement via sensory inputs is given in Fig. 22
              including feedback and feed-forward loops (Wolpert et al., 2013).
                 Contrary to negative loops and feed-forward controllers able to counter-
              act changes of physiological variables from their target values, positive feed-
              back loops amplify their initiating stimuli and move the system away from its
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