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28      PART I • Framework of Climate Science


                                                               Sediments rich in CaCO fossils occur in open-ocean
                                                                                   3
                                                            waters at depths above 3500–4000 meters (Figure 2–15).
                                                            Below that level, corrosive bottom waters dissolve calcite
                                                            shells. SiO -shelled diatoms inhabit deltas and other
                                                                     2
                                                            coastal areas and extract silica from river water flowing
                                                            off the land, but their abundance along the coasts is
                                                            masked by the influx of mud eroded from the land.
                                                            Radiolaria and diatoms are abundant in Antarctic and
                                                            equatorial regions where highly productive waters upwell
                                                            from below.
                                       Spruce                  Plankton and pollen share traits that make them
                                                            especially useful as climate proxies. Both are widely dis-
                                                            tributed: plankton live in all oceans, and pollen are
                                                            produced everywhere on continents except under ice
                                                            sheets. Also, because fossil remains of these two groups
                                                            are so abundant in sediments (usually thousands in a
                                                            tablespoon-sized sample), their relative abundances can
              60 μm                       Sage              be determined with a much higher degree of accuracy
                                                            than those fossil types that show up only sporadically.
                                                            Populations of plankton and pollen in different areas
                                                            also tend to be dominated by a small number of species
                                               Sunflower
                                Oak                         with well-defined climate preferences. The only other
                                                            organisms with comparable ranges and abundance are
        FIGURE 2-13 Pollen: a proxy indicator of climate on land  insects, which rarely leave fossil remains.
        For younger intervals, climate on land can be reconstructed
        from changes in the relative abundance of distinctive types of
        pollen. For scale, small grains of sand are 60 μm or larger in
        diameter. (Courtesy of Alan Solomon, Environmental Protection
        Agency, Corvallis, OR.)



        where they are preserved in oxygen-poor waters. Pollen
        can be identified initially by major vegetation type
        (trees, grass, and shrubs) and then further subdivided
        (spruce trees indicate cold climates; oak trees indicate
        warmth). Larger remains of vegetation that cannot have           200 μm         20 μm
        been carried far from their points of origin are also
        examined to make sure that the pollen in a lake sequence
        is representative of the nearby vegetation. These larger
        macrofossils include cones, seeds, and leaves.
           In the oceans, four major groups of shell-forming
        animal and plant plankton are used for climate recon-
        structions (Figure 2–14). Two groups form shells made
        of calcite (CaCO ). Globular sand-sized animals called
                       3                                                                  200 μm
        planktic foraminifera (upper left) inhabit the upper              2 μm
        layers of the ocean. Small spherical algae called Coccol-
        ithophoridae secrete tiny plates called  coccoliths  FIGURE 2-14 Plankton: a proxy indicator of climate in the
        (lower left) in sunlit waters. Two other groups of hard-  ocean Four types of shelled remains of plankton are common in
        shelled plankton secrete shells of opaline silica (SiO ·
                                                     2      ocean sediments: CaCO shells are represented by sand-sized
                                                                             3
        H O) and tend to thrive in productive, nutrient-rich
          2                                                 planktic foraminifera (upper left) and small clay-sized coccoliths
        surface waters.  Diatoms (upper right) are silt-sized  (lower left); SiO shells include silt-sized diatoms (upper right)
                                                                       2
        plant plankton usually shaped like either pillboxes or  and sand-sized radiolaria (lower right). For scale, small grains of
        needles. Radiolaria (lower right) are sand-sized animals  sand are 60 μm or larger in diameter. (Modified from W. F.
        with ornate shells often shaped like premodern military  Ruddiman, “Climate Studies in Ocean Cores,” in Paleoclimate Analysis
        helmets.                                            and Modeling, ed. A. D. Hecht [New York: John Wiley, 1977].)
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