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IMPLICATIONS OF PLATE TECTONICS  409



            and northwest Africa, as a consequence of the fi rst   benthic microfossils (Shackleton & Kennett, 1975), and

            phase of rifting of the supercontinent. This was initi-  the first major build-up of ice on Antarctica, coincided
            ated about 180 Ma ago. Thus the “Tethyan embayment”   with these developments, and appear to mark a transi-
            in Pangea (Fig. 13.2) was extended to the west to facili-  tion from a Greenhouse to an Icehouse Earth. The
            tate a circum-global equatorial current. This meant that   change in oxygen isotope values is particularly pro-
            some tropical waters were heated to a higher tempera-  nounced and well documented, and is essentially coin-
            ture before turning northwards and southwards to   cident with the Eocene–Oligocene boundary (Fig. 13.8).
            warm higher latitudes. In this way the whole Earth   This is also the time of the opening of the gateway
            became warmer and the temperature gradient from the   south of Tasmania (Exon et al., 2002). The full opening
            equator to the poles was further reduced.    of the Drake Passage is less well constrained, but was
               The separation of Antarctica from Africa, which   probably shortly after this (Livermore  et al., 2004).

            started about 165 Ma ago, was the first stage in the   Oxygen isotope ratios and a drop in sea level of 40 m
            break-up of Gondwana (Fig. 13.4). This was followed   suggest that during the early Oligocene the volume of
            at about 125 Ma by the rifting apart of South America   ice in Antarctica built up to perhaps as much as one-half
            and Africa, which started in the south and propagated   of its present volume. This and subsequent increases in
            northwards. This, coupled with the complex fracture   ice volume, and changes in sea level, gave rise to an
            zone pattern in the equatorial Atlantic region, due to   emergence of land areas, and a major reduction in the
            transform faulting, meant that the gateway between   area of shallow seas on continental crust (cf. Figs 13.6,
            the North and South Atlantic did not open up until   13.7).
            about 95 Ma (Fig. 13.5) (Poulsen  et al., 2001). The   Following a period of warming and deglaciation in
            initial changes in the deep-water circulation, resulting   the late Oligocene (Fig. 13.8), additional major increases
            from the opening of this gateway, may explain the   in the volume of ice on Antarctica, and associated drops
            “anoxic event” that produced the widespread black   in sea level, are thought to have occurred in the mid-
            shales in adjacent areas at that time (Poulsen  et al.,   Miocene and at the end of the Miocene. The drop in
            2001). By 95 Ma India had separated from Antarctica   sea level associated with the increase in ice volume at
            and a major Southern Ocean was opening up south   6 Ma may explain the isolation, and subsequent desicca-
            of Africa and India. However, in the late Cretaceous,   tion, of the Mediterranean Sea, as a result of the expo-
            and even in the early Cenozoic (Fig. 13.6), the circum-  sure of the sill at the Strait of Gibraltar (Van Couvering

            equatorial current still existed, and the surface water   et al., 1976), and would have restricted the flow of water
            in the high latitude oceans was still very much warmer   through the ocean gateway between North and South
            than it is today.                            America. However, additional tectonic movements
               Throughout the Cenozoic, Africa, India, and Austra-  were required before a complete land bridge formed,
            lia continued to drift northwards, away from Antarctica,   about 3 Ma ago; as determined from the interchange of
            thereby enlarging the southern and Indian Oceans, and   mammals between North and South America (Mar-
            ultimately forming the Alps and the Himalayas as a   shall, 1988). The gradual formation of the Isthmus of

            result of the collision of Africa and India with Eurasia   Panama would have led to the intensification of the
            (Section 10.4.1). By 30 Ma (Fig. 13.7), the Tethyan   Gulf Stream, and ultimately, perhaps, to the formation
            seaway was effectively closed, and the Southern Ocean   of the northern hemisphere ice-sheets (Haug &
            completely encircled Antarctica, as a result of the   Tiederman, 1998).
            opening of gateways south of Tasmania, and in the   The warm waters of the Gulf Stream would have
            Drake Passage, south of South America. These results   given rise to more warm and moist air, and hence more
            of continental drift, gave rise to major changes in the   precipitation, in relatively high latitudes in the North
            near surface oceanic circulation. There was no longer a   Atlantic area. The geographic distribution of ice sheets
            complete circum-equatorial current, and a pronounced   is determined not only by cold ambient temperatures,
            circum-polar current was established in the Southern   but also by the availability of precipitation. The Plio-
            Ocean. Thus the equatorial water became less warm,   Pleistocene ice-sheets of the northern hemisphere were
            and Antarctica was insulated from the warmer water   restricted to Greenland, northern North America, and
            circulating in the major southern hemisphere gyres of   northwest Europe for this reason. Similarly, the occur-

            the Pacific, Atlantic, and Indian Oceans. A change in   rence of tropical rain forests is determined not only
            oxygen isotope ratios in the tests of planktonic and   by high temperatures, but also by the delivery of
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