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64                              Handbook of Properties of Textile and Technical Fibres


                             High-S
                             proteins               Epicuticle
                                            Nuclear   Exocuticle
                             High-tyr       remnant  a b  Endocuticle
                             proteins
                       Low-S                                    Cuticle
                    proteins









                   Left-                    cell
                   handed              membrane                    Root end
                   coiled-coil  Matrix   complex
                   rope
           Right-                        Paracortical      Orthocortical
          handed       Microfibril  Macrofibril  cell      cell
          α- helix    (intermediate                 Mesocortical cell
                       filament)
                                                        Cortex
               1   2      7        200          2000         20,000 nm
         Figure 3.5 Schematic diagram of the structure of a fine Merino wool fiber.
         Image courtesy: CSIRO Science Images (http://www.scienceimage.csiro.au).


         a macrofibril. Several macrofibrils make up each cortical cell. Cortical cells account for
         almost 90% of the fiber bulk and are largely responsible for the mechanical properties.
         The overlapping cells are spindle shaped and approximately 100 mm long and 3e6 mm
         wide (Fig. 3.7, left), with the number and size of cells increasing with fiber diameter
         (Deng et al., 2009). They are separated by the cell membrane complex (CMC) that is



















         Figure 3.6 Transmission electron microscope image of a fine Merino wool fiber cross section
         showing the hexagonal packing of aligned intermediate filaments in the paracortex and the
         whorl-like arrangement in the orthocortex. Separating the cells is the cell membrane complex
         (CMC).
         Image courtesy of Jones LN, CSIRO.
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