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222     PART IV • Deglacial Climate Change


                     Distribution of elm pollen             forest in the north near the ice sheets and grassy steppe
                                                            vegetation prevailing farther to the south and east.
                                                            One reason for this harsh glacial climate was the clock-
                                                            wise outflow of cold winds from the Scandinavian ice
                                                            sheet (see Figure 12–11B). A second reason was the
                                                            large chilling of the North Atlantic Ocean, which
                                                            removed its moderating influence on winters in Europe
                                                            (see Figure 11–3).
                                                               One of the most striking features of the last glacial
                                                            maximum was the vast extent of steppe and tundra that
                                                            covered much of northern Asia (Figure 12–16). A region
                                                            covered today by forests of larch, birch, and alder trees
         A  Glacial observed     B  Glacial simulated       was at that time a treeless expanse of grasses and herbs.
                  <1%    1–2%     2–5%     >5%

        FIGURE 12-14 Data-model mismatch in the southeastern
        United States (A) Observed abundances of warm-adapted
        deciduous pollen such as elm in the southeastern United
        States during the glacial maximum are smaller than (B) the
        amounts simulated by climate models. (Adapted from T. Webb
        III et al., “Late Quaternary Climate Change in Eastern North
        America: A Comparison of Pollen-Derived Estimates with Climate
        Model Results,” Quaternary Science Reviews 17 [1998]: 587–606.)


           This mismatch suggests that the model-simulated
        cooling for the southeastern United States underrepre-
        sents the cooling that actually occurred by permitting too
        many warm-adapted trees. One cause of this mismatch
        may have been an unusual geographic configuration in
        which cold meltwater from the southern margin of the
        great Laurentide ice sheet flowed down the Mississippi
        River and emptied directly into the Gulf of Mexico at  A  Modern vegetation
        subtropical and tropical latitudes. If the sea-surface  Ice        Boreal forest  Mediterranean scrub
        boundary conditions used in the model had incorporated  Tundra and   Deciduous    Prairie-steppe
                                                              mountain     and conifer forest
        this cold inflow, the model might have simulated cooler
        temperatures across a broad region of the southeastern
        United States influenced by air masses from the nearby
        Gulf. Disagreements like these in initial data-model
        comparisons point the way toward future improvements
        in model simulations and data interpretation.
           Changes in Eurasia Europe was completely trans-
        formed at the glacial maximum. The conifer and decid-
        uous forests typical of today’s interglacial climate
        (Figure 12–15A) were absent from most of Europe
        south of the Scandinavian ice sheet. In their place,
        grass-covered steppes and herb-covered tundra vegeta-
        tion covered much of the continent, with bits of forest
        scattered in the south (Figure 12–15B). The moderate  B  Glacial vegetation
        maritime climate of today was preceded by a far harsher
                                                            FIGURE 12-15 Glacial north-central Europe was treeless
        continental climate, more like that of modern northern  (A) Vegetation in modern Europe is dominated by forest, with
        Asia. These differences in vegetation agree with the dry,  conifers in the north and deciduous trees to the south. (B) At
        windy conditions indicated by the greater prevalence of  the glacial maximum, Arctic tundra covered a large area south
        windblown loess (see Figure 12–4).                  of the ice sheet, with grassy steppe farther south and east and
           Biome models simulate glacial vegetation in Europe  patchy forests near the Mediterranean coasts. (Adapted from
        similar to that observed, with Arctic tundra instead of  R. F. Flint, Glacial and Quaternary Geology [New York: Wiley, 1971].)
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